For adapting to the environment, organisms have two choices: to change their physiology or to change their behavior, through genetic changes or learning; it is a trade between mutation and innovation. Innovation can be cultural if it propagates to other individuals or groups, maintains along the generations, beyond that of its discoverer and presents a certain degree of stability in its execution by various "users".
We tend to see the apes as having culture, because they are so close to us, ...
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